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Greenwood, Peter Humphry. 1980. "Towards a phyletic classification of the 'genus' Haplochromis (Pisces, Cichlidae) and related taxa. Part II; the species from lakes Victoria, Nabugabo, Edward, George and Kivu". Bulletin of the British Museum (Natural History) Zoology Series. v. 39(n. 1), pp. 1-101 (abstract)
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