mature females having the unique combination of intense dark pigmentation continuous along base of dorsal fin and on ventral surfaces from gular region to anal-fin base; and mature males having a coarse, irregular pattern of dark spots and vermiculations on cheek and opercular series, and sides with 10-12 dark stripes, each stripe occupying proximal limits of adjacent scale rows and separated by paler region central to each scale.
Ref.: http://www.pfeil-verlag.de/04biol/pdf/ief25_3_06.pdf
